Faculty of Natural
Comenius University Bratislava



Rataj Križanová F., Vďačný P. (2024) A Heterolepidoderma and Halichaetoderma gen. nov. (Gastrotricha: Chaetonotidae) riddle: integrative taxonomy and phylogeny of six new freshwater species from Central Europe. Zoological Journal of the Linnean Society 200 (2): 283–335.

Rurik I., Melichárková A., Gbúrová Štubová E., Kučera J., Kochjarová J., Paun O., Vďačný P., Slovák M. (2024). Homoplastic versus xenoplastic evolution: exploring the emergence of key intrinsic and extrinsic traits in the montane genus Soldanella (Primulaceae). The Plant Journal https://doi.org/10.1111/tpj.16630.

Zhang T., Vďačný P. (2024). Deciphering phylogenetic relationships of and delimiting species boundaries within the controversial ciliate genus Conchophthirus using an integrative morpho-evo approach. Molecular Phylogenetics and Evolution 190: e107931.


Obert T., Zhang T., Vďačný P. (2023) The search finds an end: the morphologically chimeric hysterocinetids belong to the subclass Hymenostomatia (Ciliophora: Oligohymenophorea). Zoological Journal of the Linnean Society 199 (1): 97–123.

Pecina L., Vďačný P. (2023) 2D approach to reconstruct the evolutionary history of clevelandellids (Ciliophora, Armophorea) inhabiting the hindgut of the Panesthiinae cockroaches. Zoologica Scripta 52 (6): 662‒687.

Slovák M., Melichárková A., Gbúrová Štubňová E., Kučera J., Mandáková T., Smyčka J., Lavergne S., Passalacqua N.G., Vďačný P., Paun O. (2023). Pervasive introgression during rapid diversification of the European mountain genus Soldanella (L.) (Primulaceae). Systematic Biology 72 (3): 491–504.

Weimer M., Vďačný P., Wolf M. (2023). Paramecium: RNA sequence–structure phylogenetics. International Journal of Systematic and Evolutionary Microbiology 73 (4): e005744.

Zhang T., Rurik I., Vďačný P. (2023). A holistic approach to inventory the diversity of mobilid ciliates (Protista: Ciliophora: Peritrichia). Organisms Diversity & Evolution 23 (3): 425–454.

Zhang T., Vďačný P. (2023) Morphological and molecular characterization of the ciliate parasite Tetrahymena rostrata infecting the renal organ of the dusky slug (Arion fuscus). Canadian Journal of Zoology 101 (1): 32–45.

Zhang T., Vďačný P. (2023). Re-discovery and novel contributions to morphology and multigene phylogeny of Protospirella mazurica (Raabe, 1968) Aescht, 2001 (Ciliophora: Pleuronematida), an obligate symbiont of the river nerite Theodoxus fluviatilis Linnaeus, 1758 (Mollusca: Gastropoda). European Journal of Protistology 88: e125956.


Kodada J., Boukal D.S., Vďačný P., Goffová K., Ondrejková K. (2022) Elmidae of Sarawak: the genus Potamophilus Germar, 1811, with a description of P. kelabitensis sp. nov. (Insecta: Coleoptera). European Journal of Taxonomy 806: 1‒18.

Obert T., Zhang T., Rurik I., Vďačný P. (2022) First molecular evidence of hybridization in endosymbiotic ciliates (Protista, Ciliophora). Frontiers in Microbiology 13: e1067315.

Pecina L., Vďačný P. (2022) DNA barcoding and coalescent-based delimitation of endosymbiotic clevelandellid ciliates (Ciliophora: Clevelandellida): a shift to molecular taxonomy in the inventory of ciliate diversity in panesthiine cockroaches. Zoological Journal of the Linnean Society 194 (4): 1072‒1102.

Rataj M., Vďačný P. (2022) Putative ITS2 secondary structure model and multi‐gene phylogenies of tetrahymenids (Ciliophora, Hymenostomatia) parasitizing planarians and crayfish worms. Zoologica Scripta 51 (2): 246‒265.

Rataj M., Zhang T., Vďačný P. (2022) Nuclear and mitochondrial SSU rRNA genes reveal hidden diversity of Haptophrya endosymbionts in freshwater planarians and challenge their traditional classification in Astomatia. Frontiers in Microbiology 13: e830951.

Rataj Križanová F., Vďačný P. (2022) A huge undescribed diversity of the subgenus Hystricochaetonotus (Gastrotricha, Chaetonotidae, Chaetonotus) in Central Europe. European Journal of Taxonomy 840: 1-93.

Zhang G., Sheng Y., Liu Y., Cao X., Al‑Farraj S.A., Vďačný P., Pan H. (2022) Integrative studies on three new freshwater Amphileptus species (Ciliophora, Pleurostomatida) discovered in northern China. Marine Life Science & Technology  4 (4): 452–470.

Zhang T., Shao C., Song W., Vďačný P., Al-Farraj S.A., Wang Y. (2022) Multi-gene phylogeny of the ciliate genus Trachelostyla (Ciliophora, Hypotrichia), with integrative description of two species, Trachelostyla multinucleata spec. nov. and T. pediculiformis (Cohn, 1866). Frontiers in Microbiology 12: e775570.

Zhang T., Vďačný P. (2022) Multiple independent losses of cell mouth in phylogenetically distant endosymbiotic lineages of oligohymenophorean ciliates: A lesson from Clausilocola. Molecular Phylogenetics and Evolution 166: e107310.


Kodada J., Selnekovič D., Jäch M.A., Goffová K., Vďačný P. (2021) Taxonomic revision of the genus Elmomorphus Sharp, 1888 I. Japanese and Korean species (Coleoptera: Dryopidae). European Journal of Taxonomy 758, 97‒121.

Križanová F., Vďačný P. (2021) Description of Lepidochaetus tirjakovae sp. nov.(Gastrotricha: Paucitubulatina: Chaetonotidae), using morphology and DNA barcoding. Zoologischer Anzeiger 292: 207‒224.

Obert T., Vďačný P. (2021) Multi-gene phylogeny of the subclass Astomatia (Protista: Ciliophora) refreshed with two rare astome ciliates from the digestive tube of endogeic earthworms. Organisms Diversity & Evolution 21 (1): 59‒77.

Obert T., Rurik I., Vďačný P. (2021) Diversity and eco-evolutionary associations of endosymbiotic astome ciliates with their lumbricid earthworm hosts. Frontiers in Microbiology 12: e689987.

Rajter Ľ., Ewers I., Graupner N., Vďačný P., Dunthorn M. (2021) Colpodean ciliate phylogeny and reference alignments for phylogenetic placements. European Journal of Protistology 77: e125747.

Rataj M., Vďačný P. (2021) Cryptic host-driven speciation of mobilid ciliates epibiotic on freshwater planarians. Molecular Phylogenetics and Evolution 161: e107174.

Vďačný P., Foissner W. (2021) Morphology and ontogenesis of two new Hemiholosticha species (Ciliophora, Hypotrichia, Hemiholostichidae nov. fam.). European Journal of Protistology 77: e125763.

Zhang T., Vďačný P. (2021) Re-discovery and novel contributions to morphology and multigene phylogeny of Myxophyllum steenstrupi (Ciliophora: Pleuronematida), an obligate symbiont of terrestrial pulmonates. Zoological Journal of the Linnean Society 192 (1): 1‒23.

Zhang T., Vďačný P. (2021) A discovery of two new Tetrahymena species parasitizing slugs and mussels: morphology and multi-gene phylogeny of T. foissneri sp. n. and T. unionis sp. n. Parasitology Research 120 (7): 2595‒2616.


Fačkovcová Z., Slovák M., Vďačný P., Melichárková A., Zozomová-Lihová J., Guttová A. (2020) Spatio-temporal formation of the genetic diversity in the Mediterranean dwelling lichen during the Neogene and Quaternary epochs. Molecular Phylogenetics and Evolution 144: e106704.

Obert T., Vďačný P. (2020) Delimitation of five astome ciliate species isolated from the digestive tube of three ecologically different groups of lumbricid earthworms, using the internal transcribed spacer region and the hypervariable D1/D2 region of the 28S rRNA gene. BMC Evolutionary Biology 20: e37.

Obert T., Vďačný P. (2020) Evolutionary origin and host range of Plagiotoma lumbrici (Ciliophora, Hypotrichia), an obligate gut symbiont of lumbricid earthworms. Journal of Eukaryotic Microbiology 67: 176‒189.

Pecina L., Vďačný P. (2020) Morphological versus molecular delimitation of ciliate species: A case study of the family Clevelandellidae (Protista, Ciliophora, Armophorea). European Journal of Taxonomy 697: 1‒46.

Pecina L., Vďačný P. (2020) Two new endozoic ciliates, Clevelandella lynni sp. n. and Nyctotherus galerus sp. n., isolated from the hindgut of the wood-feeding cockroach Panesthia angustipennis (Illiger, 1801). Journal of Eukaryotic Microbiology 67: 436‒449.

Rataj M., Vďačný P. (2020) Multi-gene phylogeny of Tetrahymena refreshed with three new histophagous species invading freshwater planarians. Parasitology Research 119: 1523‒1545.

Zhang T., Wang Y., Cheng T., Ma J., Vďačný P., Song W., Shao C. (2020) Systematics and multi-gene phylogeny of the subfamily Nothoholostichinae (Ciliophora, Hypotrichia), with integrative description of a new marine species Nothoholosticha luporinii n. sp. Frontiers in Marine Science 7: e610886.


Bokor B., Soukup M., Vaculík M., Vďačný P., Weidinger M., Lichtscheidl I., Vávrová S., Šoltys K., Sonah H., Deshmukh R., Bélanger R.R., White P.J., El-Serehy H.A., Lux A. (2019) Silicon uptake and localisation in date palm (Phoenix dactylifera)–a unique association with sclerenchyma. Frontiers in Plant Science 10: e988.

Obert T., Vďačný P. (2019) Integrative taxonomy of five astome ciliates (Ciliophora, Astomatia) isolated from earthworms in Central Europe. European Journal of Taxonomy 559: 1–37.

Rajter Ľ., Bourland W., Vďačný P. (2019) Morpho-molecular characterization of the litostomatean predatory ciliate Phialina pupula (Müller, 1773) Foissner, 1983 (Haptoria, Lacrymariidae). Acta Protozoologica 58: 53–68.

Rataj M., Vďačný P. (2019) Living morphology and molecular phylogeny of oligohymenophorean ciliates associated with freshwater turbellarians. Diseases of Aquatic Organisms 134: 147–166.

Shazib S.U.A., Vďačný P., Slovák M., Gentekaki E., Shin M.K. (2019) Deciphering phylogenetic relationships and delimiting species boundaries using a Bayesian coalescent approach in protists: A case study of the ciliate genus Spirostomum (Ciliophora, Heterotrichea). Scientific Reports 9: e16360.

Vďačný P., Foissner W. (2019) Re-analysis of the 18S rRNA gene phylogeny of the ciliate class Colpodea. European Journal of Protistology 67: 89–105.

Vďačný P., Foissner W. (2019) A huge diversity of metopids (Ciliophora, Armophorea) in soil from the Murray River floodplain, Australia. III. Morphology, ontogenesis and conjugation of Metopus boletus nov. spec., with implications for the phylogeny of the SAL supercluster. European Journal of Protistology 69: 117–137.

Vďačný P., Foissner W. (2019) Morphology and ontogenesis of Hemiholosticha pantanalensis nov. spec. (Ciliophora, Hypotrichia, Psilotrichidae). Acta Protozoologica 58: 93‒113.

Vďačný P., Rajter Ľ., Stoeck T., Foissner W. (2019) A proposed timescale for the evolution of armophorean ciliates: clevelandellids diversify more rapidly than metopids. Journal of Eukaryotic Microbiology 66: 167–181.



Bohatová M., Vďačný P. (2018) Locomotory behaviour of two phylogenetically distant predatory ciliates: does evolutionary history matter? Ethology Ecology & Evolution 30: 195–219.

Košťál M., Vďačný P. (2018) On the phylogenetic position of the weevil tribe Acentrusini Alonso-Zarazaga, 2005 (Coleoptera: Curculionidae: Curculioninae). Diversity 10: e34.

Rajter Ľ., Vďačný P. (2018) Selection and paucity of phylogenetic signal challenge the utility of alpha-tubulin in reconstruction of evolutionary history of free-living litostomateans (Protista, Ciliophora). Molecular Phylogenetics and Evolution 127: 534–544.

Rataj M., Vďačný P. (2018) Dawn of astome ciliates in light of morphology and time-calibrated phylogeny of Haptophrya planariarum, an obligate endosymbiont of freshwater turbellarians. European Journal of Protistology 64: 54–71.

Slovák M., Kučera J., Lack H.W., Ziffer-Berger J., Melicharková A., Záveská E., Vďačný P. (2018). Diversification dynamics and transoceanic Eurasian-Australian disjunction in the genus Picris (Compositae) induced by the interplay of shifts in intrinsic/extrinsic traits and paleoclimatic oscillations. Molecular Phylogenetics and Evolution 119: 182–195.

Vďačný P. (2018) Evolutionary associations of endosymbiotic ciliates shed light on the timing of the marsupial-placental split. Molecular Biology and Evolution 35: 1757–1769.

Vďačný P., Érseková E., Šoltys K., Budiš J., Pecina L., Rurik I. (2018) Co-existence of multiple bacterivorous clevelandellid ciliate species in hindgut of wood-feeding cockroaches in light of their prokaryotic consortium. Scientific Reports 8: e17749.

Vršanský P., Bechly G., Zhang Q., Jarzembowski E.A., Mlynský T., Šmídová L., Barna P., Kúdela M., Aristov D., Bigalk S., Krogmann L., Li L., Zhang Q., Zhang H., Ellenberger S., Müller P., Gröhn C., Xia F., Ueda K., Vďačný P., Valaška D., Vršanská L., Wang B. (2018) Batesian insect-insect mimicry-related explosive radiation of ancient alienopterid cockroaches. Biologia (Bratislava) 73: 987–1006.





Jang S.W., Vďačný P., Shazib S.U.A., Shin M.K. (2017) Linking morphology and molecules: integrative taxonomy of spathidiids (Protista: Ciliophora: Litostomatea) from Korea. Journal of Natural History 51: 939–974.

Rajter Ľ., Vďačný P. (2017) Constraints on phylogenetic interrelationships among four free-living litostomatean lineages inferred from 18S rRNA gene-ITS region sequences and secondary structure of the ITS2 molecule. Acta Protozoologica 56: 255–281.

Vďačný P. (2017) Integrative taxonomy of ciliates: Assessment of molecular phylogenetic content and morphological homology testing. European Journal of Protistology 61: 388–398.

Vďačný P., Foissner W. (2017) A huge diversity of metopids (Ciliophora, Armophorea) in soil from the Murray River floodplain, Australia. I. Description of five new species and redescription of Metopus setosus Kahl, 1927. European Journal of Protistology 57: 35–76.

Vďačný P., Foissner W. (2017) A huge diversity of metopids (Ciliophora, Armophorea) in soil from the Murray river floodplain, Australia. II. Morphology and morphogenesis of Lepidometopus platycephalus nov. gen., nov. spec. Acta Protozoologica 56: 39–57.

Vďačný P., Rajter Ľ., Shazib S.U.A., Jang S.W., Shin M.K. (2017) Diversification dynamics of rhynchostomatian ciliates: the impact of seven intrinsic traits on speciation and extinction in a microbial group. Scientific Reports 7: e9918.

Vďačný P., Rataj M. (2017) Evaluation of systematic position of helicoprorodontids and chaeneids (Ciliophora, Litostomatea): an attempt to break long branches in 18S rRNA gene phylogenies. Journal of Eukaryotic Microbiology 64: 608–621.




Benčaťová S., Tirjaková E., Vďačný P. (2016) Resting cysts of Parentocirrus hortualis Voß, 1997 (Ciliophora, Hypotrichia), with preliminary notes on encystation and various types of excystation. European Journal of Protistology 53: 45–60.

Petrýdesová J., Kučera J., Bacigálová K., Vadkertiová R., Lopandic K., Vďačný P., Slovák M. (2016) Disentangling identity of Taphrina species parasitizing herbaceous Rosaceae, with proposal of T. gei-montani sp. nov. International Journal of Systematic and Evolutionary Microbiology 66: 2540–2549.

Rajter Ľ., Vďačný P. (2016) Rapid radiation, gradual extinction and parallel evolution challenge generic classification of spathidiid ciliates (Protista, Ciliophora). Zoologica Scripta 45: 200–223.

Shazib S.U.A., Vďačný P., Kim J.H., Jang S.W., Shin M.K. (2016) Molecular phylogeny and species delimitation within the ciliate genus Spirostomum (Ciliophora, Postciliodesmatophora, Heterotrichea), using the internal transcribed spacer region. Molecular Phylogenetics and Evolution 102: 128–144.

Tirjaková E., Krajčovičová K., Illyová M., Vďačný P. (2016) Interaction of ciliate communities with cyanobacterial water bloom in a shallow, hypertrophic reservoir. Acta Protozoologica 55: 173–188.


Dunthorn M., Lipps J.H., Dolan J.R.,  Abboud-Abi Saab M., Aescht E., Bachy C., Barría de Cao, M.S., Berger H., Bourland W.A., Choi J.K., Clamp J., Doherty M., Gao F., Gentekaki E., Gong J., Hu X., Huang J., Kamiyama T., Johnson M.D., Kammerlander B., Kim S.Y., Kim Y.-O., la Terza A., Laval-Peuto M., Lipscomb D., Lobban C.S., Long H., Luporini P., Lynn D.H., Macek M., Mansergh R.I., Martín-Cereceda M., McManus G.G., Montagnes D.J.S., Ong'ondo G.O., Patterson D.J., Pérez-Uz B., Quintela-Alonso P., Safi L.S.L., Santoferrara L.F., Sonntag B., Song W., Stoeck T., Stoecker D.K., Strüder-Kypke M.C., Trautmann I., Utz L.R.P., Vallesi A., Vďačný P., Warren A., Weisse T., Wickham S.A., Yi Z., Zhang W., Zhan Z., Zufall R., Agatha S. (2015) Ciliates — Protists with complex morphologies and ambiguous early fossil record. Marine Micropaleontology 119: 1–6.

Jang S.W., Vďačný P., Shazib S.U.A., Shin M.K. (2015) Morphology, ciliary pattern and molecular phylogeny of Trachelophyllum brachypharynx Levander, 1894 (Litostomatea, Haptoria, Spathidiida). Acta Protozoologica 54: 123–135.

Tirjaková E., Botlíková S., Vďačný P. (2015) Checklist and and distribution of ciliates from the family Euplotidae Ehrenberg, 1838 (Protista: Ciliophora: Spirotrichea) in Sovakia, Central Europe. Zootaxa 3920: 343–365.

Vďačný P. (2015) Estimation of divergence times in litostomatean ciliates (Ciliophora: Intramacronucleata), using Bayesian relaxed clock and 18S rRNA gene. European Journal of Protistology 51: 321–334.

Vďačný P., Rajter Ľ. (2015)  Reconciling morphological and molecular classification of predatory ciliates: Evolutionary taxonomy of dileptids (Ciliophora, Litostomatea, Rhynchostomatia). Molecular Phylogenetics and Evolution 90: 112–128.

Vďačný P., Rajter Ľ., Shazib S.U., Jang S.W., Kim J.H., Shin M.K. (2015) Reconstruction of evolutionary history of pleurostomatid ciliates (Ciliophora, Litostomatea, Haptoria): interplay of morphology and molecules. Acta Protozoologica 54: 9–29.


Bourland W.A., Wendell L., Hampikian G., Vďačný P. (2014) Morphology and phylogeny of Bryophryoides ocellatus n. g., n. sp. (Ciliophora, Colpodea) from in situ soil percolates of Idaho, U.S.A. European Journal of Protistology 50: 47–67.

Jang S.W., Vďačný, P., Shazib S.U.A., Shin M.K. (2014) Morphological and molecular characterization of the name-bearing type species Rimaleptus binucleatus (Kahl, 1931), with a phylogenetic re-analysis of dileptid evolutionary history (Ciliophora: Litostomatea: Rhynchostomatia). European Journal of Protistology 50: 456–471.

Kim J.H., Vďačný P., Shazib S.U.A., Shin M.K. (2014) Morphology and molecular phylogeny of Apoterritricha lutea n. g., n. sp (Ciliophora, Spirotrichea, Hypotrichia): A putative missing link connecting Cyrtohymena and Afrokeronopsis. Journal of Eukaryotic Microbiology 61: 520–536.

Kwon C.B., Vďačný P., Shazib S.U.A., Shin M.K. (2014) Morphology and molecular phylogeny of a new haptorian ciliate, Chaenea mirabilis sp n., with implications for the evolution of the dorsal brush in haptorians (Ciliophora, Litostomatea). Journal of Eukaryotic Microbiology 61: 278–292.

Shazib S.U.A., Vďačný P., Kim J.H., Jang S.W., Shin M.K. (2014) Phylogenetic relationships of the ciliate class Heterotrichea (Protista, Ciliophora, Postciliodesmatophora) inferred from multiple molecular markers and multifaceted analysis strategy. Molecular Phylogenetics and Evolution 78: 118–135.

Slovák M., Kučera J., Záveská E., Vďačný P. (2014) Dealing with discordant genetic signal caused by hybridisation, incomplete lineage sorting and paucity of primary nucleotide homologies: A case study of closely related members of the genus Picris subsection Hieracioides (Compositae). Plos One 9: e104929.

Vďačný P., Breiner H.-W., Yashchenko V., Dunthorn M., Stoeck T., Foissner W. (2014) The chaos prevails: molecular phylogeny of the Haptoria (Ciliophora, Litostomatea). Protist 165: 93–111.

Vďačný P., Rajter Ľ. (2014) An annotated and revised checklist of pleurostome ciliates (Protista: Ciliophora: Litostomatea) from Slovakia, Central Europe. Zootaxa 3760: 501–521.

Vďačný P., Slovák M., Foissner W. (2014) Multivariate morphometric analyses of the predatory ciliate genus Semispathidium (Ciliophora: Litostomatea), with description of S. longiarmatum nov. spec. European Journal of Protistology 50: 329–344.


Tirjaková E., Vďačný P. (2013) Recovery of ciliate communities from an oligotrophic mountain stream after a catastrophic wind damage. European Journal of Protistology 49: 526–537.

Tirjaková E., Vďačný P. (2013) Analysis and evolution of water quality of the upper Váh River (northern Slovakia) by long-term changes in the community structure of ciliates (Protista: Ciliophora). Biologia (Bratislava) 68: 667–678.

Vďačný P., Foissner W. (2013) Synergistic effects of combining morphological and molecular data in resolving the phylogenetic position of Semispathidium (Ciliophora, Haptoria) with description of Semispathidium breviarmatum sp. n. from tropical Africa. Zoologica Scripta 42: 529–549.


Bourland W.A., Hampikian G., Vďačný P. (2012) Morphology and phylogeny of a new woodruffiid ciliate, Etoschophrya inornata sp. n. (Ciliophora, Colpodea, Platyophryida), with an account on evolution of platyophryids. Zoologica Scripta 41: 400–416.

Holecová M., Christophoryová J., Mrva M., Roháčová M., Stašiov S., Štrichelová J., Šustek Z., Tirjaková E., Tuf I.H., Vďačný P., Zlinská J. (2012) Biodiversity of soil micro- and macrofauna in oak-hornbeam forest ecosystem on the territory of Bratislava. Comenius University: Bratislava.

Orsi W., Charver S., Vďačný P., Bernhard J.M., Edgcomb V.P. (2012) Prevalence of partnerships between bacteria and ciliates in oxygen-depleted marine water columns. Frontiers in Microbiology 3: e341.

Orsi W., Edgcomb V., Faria J., Foissner W., Fowle W.H., Hohmann T., Suarez P., Taylor C., Taylor G.T., Vďačný P., Epstein S.S. (2012) Class Cariacotrichea, a novel ciliate taxon from the anoxic Cariaco Basin, Venezuela. International Journal of Systematic and Evolutionary Microbiology 62: 1425–1433.

Vďačný P., Bourland W.A., Orsi W., Epstein S.S., Foissner W. (2012) Genealogical analyses of multiple loci of litostomatean ciliates (Protista, Ciliophora, Litostomatea). Molecular Phylogenetics and Evolution 65: 397–411.

Vďačný P., Foissner W. (2012) Monograph of the dileptids (Protista, Ciliophora, Rhynchostomatia). Denisia 31: 1–529.

Vďačný P., Tirjaková E. (2012) Taxonomic revision and neotypification of Zosterodasys transversus (Kahl, 1928), with description of a mirror-image doublet (Ciliophora, Phyllopharyngea, Synhymeniida). Acta Protozoologica 51: 39–54.

Vďačný P., Tirjaková E. (2012) Taxonomic revision of the ciliate genus Zosterodasys Deroux, 1978 (Protista: Ciliophora: Synhymeniida). Zootaxa 3345: 34–58.


Bourland W.A., Vďačný P., Davis M.C., Hampikian G. (2011) Morphology, morphometrics, and molecular characterization of Bryophrya gemmea n. sp. (Ciliophora, Colpodea): Implications for the phylogeny and evolutionary scenario for the formation of oral ciliature in the order Colpodida. Journal of Eukaryotic Microbiology 58: 22–36.

Edgcomb V., Orsi W., Taylor G.T., Vdacny P., Taylor C., Suarez P., Epstein S. (2011) Accessing marine protists from the anoxic Cariaco Basin. ISME Journal 5: 1237–1241.

Vďačný P., Bourland W.A., Orsi W., Epstein S.S., Foissner W. (2011) Phylogeny and classification of the Litostomatea (Protista, Ciliophora), with emphasis on free-living taxa and the 18S rRNA gene. Molecular Phylogenetics and Evolution 59: 510–522.

Vďačný P., Orsi W., Bourland W.A., Shimano S., Epstein S.S., Foissner W. (2011) Morphological and molecular phylogeny of dileptid and and tracheliid ciliates: Resolution at the base of the class Litostomatea (Ciliophora, Rhynchostomatia). European Journal of Protistology 47: 295–313.


Vďačný P., Orsi W., Foissner W. (2010) Molecular and morphological evidence for a sister group relationship of the classes Armophorea and Litostomatea (Ciliophora, Intramacronucleata, Lamellicorticata infraphyl. nov.), with an account on basal haptorid litostomateans. European Journal of Protistology 46: 298–309.

Vďačný P., Tirjaková E., Tóthová T., Pristaš P., Javorský P. (2010) Morphological and phylogenetical studies on a new soil hypotrich ciliate: Kahliella matisi spec. nov. (Hypotrichia, Kahliellidae). European Journal of Protistology 46: 319–333.


Vďačný P., Foissner W. (2009) Ontogenesis of Dileptus terrenus and Pseudomonilicaryon brachyproboscis (Ciliophora, Haptoria). Journal of Eukaryotic Microbiology 56: 232–243.


Vďačný P., Foissner W. (2008) Morphology, conjugation, and postconjugational reorganization of Dileptus tirjakovae n. sp. (Ciliophora, Haptoria). Journal of Eukaryotic Microbiology 55: 436–447.

Vďačný P., Foissner W. (2008) Description of four new soil dileptids (Ciliophora, Haptoria), with notes on adaptations to the soil environment. Acta Protozoologica 47: 211–230.


Vďačný P. (2007) Morphological and taxonomical studies on two soil haptorid ciliates (Ciliophora, Litostomatea): Clavoplites haranti sp. n. and Enchelys terrenum (Foissner, 1984) comb. n., and taxonomy of the family Enchelyidae Ehrenberg, 1838. European Journal of Protistology 43: 225–237.

Vďačný P. (2007) Morphological variation of Fuscheria terricola Berger et al., 1983 (Ciliophora, Haptoria). Linzer biologische Beiträge 39: 1239–1248.

Vďačný P. (2007) Morphology and infraciliature of the soil ciliate Metopus hasei Sondheim, 1929 (Ciliophora, Armophorida) from Biele Karpaty Mountains (Slovakia). Protistology 5: 231–236.

Vďačný P., Tirjaková E. (2007) Observations on some soil haptorid ciliates (Protozoa, Ciliophora) from Slovakia. Biologia (Bratislava) 62: 720–730.


Vďačný P., Tirjaková E. (2006) A new soil hypotrich ciliate (Protozoa, Ciliophora) from Slovakia: Gonostomum albicarpathicum nov. spec. European Journal of Protistology 42: 91–96.

Vďačný P., Hlúbiková D., Tirjaková E. (2006) Spathidium seppelti foissneri nov. subspec., Spathidium simplinucleatum nov. stat., and Dileptus americanus Kahl, 1931, one new and two poorly known gymnostome ciliates from soils of Slovakia. European Journal of Protistology 42: 175–189.

Vďačný P., Tirjaková E.  (2006) The first record of soil ciliates (Protozoa, Ciliophora) from classes Prostomea, Nassophorea, Spirotrichea, and Colpodea in Slovakia. Biologia (Bratislava) 61: 509–516.


Tirjaková E., Vďačný P. (2005) Ciliate communities (Protozoa, Ciliophora) in tree-holes and influence of selected environmental factors on their structure. Ekológia (Bratislava) 25 (Supplement 2): 20-36.


Tirjaková E., Vďačný P. (2004) Prvé nálezy druhov Holostichides chardezi Foissner, 1987 a Australocirrus octonucleatus Foissner, 1988 (Ciliophora: Hypotrichia) na území Slovenska [First records of two species of ciliates: Holostichides chardezi and Australocirrus octonucleatus (Ciliophora: Hypotrichia) in Slovakia]. Folia faunistica Slovaca 9: 25–30.